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KOLREUTER'S CONTRIBUTIONS TO BIOLOGY 155
Table 1. Kolreuter's intrageneric crosses as reported in the Vorlaufige Nachricht
F, crosses: Simple hybridizations of species
Nicotiana rustica P x N. paniculata 6 (A:39-41, B:23-24, C:90)
Nicotiana paniculata 9 x N. rustica 6 (A:43, B: 10-1 1, C:90)
Nicotiana major vulgaris P x N. glutinosa 6 (B:27-30, C:90)
Nicotiana transylvanica P x N. glutinosa 6 (B:31, C:90)
Nicotiana rustica P x N. perennis 8 (B:64)
Nicotiana perennis P x N. rustica 6 (B:64)
Nicotiana rustica P x N. major 6 (B:64)
Nicotiana major P x N. rustica 6 (B:64)
Nicotiana major var. P x N. rustica 6 (B:64)
Nicotiana rustica P x N. glutinosa 6 (B:64-65)
Nicotiana glutinosa P x N. rustica 8 (B:65)
Nicotiana paniculata P x N. perennis 8 (B:65)
Nicotiana perennis P x N. paniculata 8 (B:65)
Nicotiana paniculata P x N. major 6 (B:66)
Nicotiana major P x N. paniculata 6 (B:66)
Nicotiana paniculata P x N. glutinosa 6 (B:66)
Nicotiana glutinosa P x N. paniculata 6 (B:66-67)
Nicotiana glutinosa P x N. major 8 (B:67, C:58-60)
Nicotiana glutinosa P x N. perennis 6 (B:67-68, C:58-60)
Nicotiana perennis P x N. glutinosa 8 (B:67-68, C:56-58)
Nicotiana majorflore albo P x N. glutinosa 6 (B:68, C:56-58)
Nicotiana paniculata P x N. glutinosa 8 (C:41-44)
Nicotiana paniculata P x N. transylvanica (C:46-48)
Nicotiana paniculata P x N. perennis 6 (C:48)
Nicotiana, five cultivars, all possible crosses (C:.53-56)
Dianthus chinensis P x D. carthusianorum 6 (B:43-45, C: 118-1 19)
Dianthus chinensis P x D. hortensis 8 (C: 108-1 12)
Dianthus chinensis P x D. carthusianorum sylv. 6 (C: 112-1 17)
Dianthus barbatus P x D. chinensis 6 (D:5.5-58)
Dianthus hortensis P x D. chinensis 6 (D:66-67)
Dianthus chinensis P x D. superbus 6 (D:69-74)
Dianthus barbatus P x D. hortensis 6 (D:77-78)
Dianthus hortensis P x D. barbatus 6 (D:79)
Dianthus barbatus P x D. deltoides 6 (D:79-82)
Dianthus chinensis P x D. deltoides 6 (D:82-84)
Dianthus chinensis P x D. Armeria 6 (D:86-89)
Dianthus plumarius Sibiricus P x D. chinensis 6 (D:89-93)
Dianthus plumarius Sibiricus P x D. glaucus 6 (D:93-96)
Dianthus various crosses (D: 112-1 14)
NOTES. The crosses in each category are arranged in order of first appearance. Many of Kolreuter's
abbreviations have been spelled out; those that remain (except for some uses of the common singleletter
abbreviation of the genus) are Kolreuter's own. Botanical nomenclature, even for well-known
species, was not yet stable in the 1760s; therefore students of Kolreuter's crosses should be warned
that, e.g., what he called Dianthus carthus [ianorum] (always abbreviated) in his earlier writings he
later (C and D)called D. barbat[us],a nd what he first called Dianthus plumar[ius] he later called D.
superbus. My own account will have to be scrutinized by botanists for the correctness of the taxonomy.
156 ERNST MAYR
Table 1--Continued
Verbascum phoeniceum Q x V. Lichnitis 6 (C:9, C: 12-18)
Verbascum phoeniceum Q x V. phlomoides 6 (C:9, C:18-24)
Verbascum phoeniceum Q x V. nigrum 6 (C: 10, C:25-30)
Verbascum phoeniceum Q x V. Blattaria 6 (C:10, C:31-36)
Verbascum phoeniceum Q x V. phoeniceum 6 selfed (C: 10-1 1 )
Verbascum phoeniceum Q x V. thapsus 6 (D:2-5)
Verbascum Lychnitisflore albo Q x V. phoeniceum 6 (D:5-6)
Verbascum nigrum Q x V. Lychnitis fl. alb. 6 (D:6-10)
Verbascum Lychnitis flore albo Q x V. nigrlrm 6 (D: 10)
Verbascum nigrum Q x V. Blattaria flore flavo (D: 11-15)
Verbascum Blattaria flore flavo Q x V. phoeniceum 6 (D: 16)
Verbascum Lychnitis flore albo Q x V. Blattaria Jlore flavo 6 (D: 17-20)
Verbascum Blattaria flore flavo Q x V. Lychnitis flore albo 6 (D:21)
Verbascum nigrum Q x V. thapsus 6 (D:21-26)
Verbascum thapsus Q x V. nigrum 6 (D:26)
Verbascum phlomoides Q x V. nigrum 6 (D:27-29)
Verbascum phlomoides Q x V. Lychnitisflore albo 6 (D:29-33)
Verbascum thapsus Q x V. Lychnitisflore albo 6 (D:33-34)
Verbascum Lychnitis flore albo Q x V. thapsus 6 (D:34)
Verbascum nigrum Q x V. Lychnitis flore Jlavo 6 (D:35)
Verbascum Blattaria flore flavo Q x V. Lychnitis flore flavo 6 (D:36)
Verbascum thapsus Q x V. Blattariafloreflavo 6 (D:46)
Verbascum Blattaria flore flavo Q x V. thapsus 6 (D:46)
Aquilegia vulgaris Q x A. canadensis 6 (D: 119-124)
Aquilegia canadensis Q x A. vulgaris 6 (D:119-124)
F, crosses
(Nicotiana rustica Q x N. paniculata 6 ) selfed (B:21)
(Dianthus chinensis Q x D. carthusianorum 6 ) selfed (B:44)
All hybrid Verbascum selfed (C:39-40)-totally sterile
(Dianthus chinensis Q x D. carthusianorum 6 ) selfed (C:106)
(D. chinensis Q x D. carthusianorum sylv. 6 ) selfed (C:117)
(Nicotiana rustica Q x N. paniculata 6 ) 2nd backcross selfed (D:53)
([N. paniculata Q x N. rustica 61 Q x N. rustica 6 ) selfed (D:53-54)
(Dianthus chinensis Q x D. hortensis 6 ) selfed (D:65-66)
(D. barbatusa Q x D, chinensis 6 ) selfed (D: 105-106)
(D. chinensis Q x D. barbatusa 6 ) selfed (D:106-107)
Simple backcrosses
(Nicotiana rustica Q x N. paniculata 6 ) Q x N. rustica 6 (B:12-15, C:90)
(N. rustica Q x N. paniculata 6 ) Q x N, paniculata 6 (B:15-18, C:90)
(N. paniculata Q x N. rustica 6 ) Q x N. paniculata 6 (C:85)
(N. paniculata Q x N. rustica 6 ) Q x N. rustica 6 (C:86-87)
a Kolreuter has Dianth. barbar., no doubt a misprint for barbat.